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Owls are regarded as having the most frontally placed eyes among all avian groups, which gives them some of the largest binocular fields of vision.

However, owls are farsighted and cannot focus on objects within a few centimeters of their eyes. Owls exhibit specialized hearing functions and ear shapes that also aid in hunting.

They are noted for asymmetrical ear placements on the skull in some genera. Owls can have either internal or external ears, both of which are asymmetrical.

Asymmetry has not been reported to extend to the middle or internal ear of the owl. Asymmetrical ear placement on the skull allows the owl to pinpoint the location of its prey.

This is especially true for strictly nocturnal species such as the barn owls Tyto or Tengmalm's owl. This time difference between ears is about 30 microseconds.

Behind the ear openings are modified, dense feathers, densely packed to form a facial ruff, which creates an anterior-facing, concave wall that cups the sound into the ear structure.

The facial disk also acts to direct sound into the ears, and a downward-facing, sharply triangular beak minimizes sound reflection away from the face.

The shape of the facial disk is adjustable at will to focus sounds more effectively. The prominences above a great horned owl's head are commonly mistaken as its ears.

This is not the case; they are merely feather tufts. The ears are on the sides of the head in the usual location in two different locations as described above.

While the auditory and visual capabilities of the owl allow it to locate and pursue its prey, the talons and beak of the owl do the final work.

The owl kills its prey using these talons to crush the skull and knead the body. The Tasmanian masked owl has some of the proportionally longest talons of any bird of prey; they appear enormous in comparison to the body when fully extended to grasp prey.

The family Tytonidae has inner and central toes of about equal length, while the family Strigidae has an inner toe that is distinctly shorter than the central one.

The beak of the owl is short, curved, and downward-facing, and typically hooked at the tip for gripping and tearing its prey.

Once prey is captured, the scissor motion of the top and lower bill is used to tear the tissue and kill. The sharp lower edge of the upper bill works in coordination with the sharp upper edge of the lower bill to deliver this motion.

The downward-facing beak allows the owl's field of vision to be clear, as well as directing sound into the ears without deflecting sound waves away from the face.

The coloration of the owl's plumage plays a key role in its ability to sit still and blend into the environment, making it nearly invisible to prey.

Owls tend to mimic the coloration and sometimes the texture patterns of their surroundings, the barn owl being an exception.

The snowy owl Bubo scandiacus appears nearly bleach-white in color with a few flecks of black, mimicking their snowy surroundings perfectly, while the speckled brown plumage of the tawny owl Strix aluco allows it to lie in wait among the deciduous woodland it prefers for its habitat.

Likewise, the mottled wood-owl Strix ocellata displays shades of brown, tan and black, making the owl nearly invisible in the surrounding trees, especially from behind.

Usually, the only tell-tale sign of a perched owl is its vocalizations or its vividly colored eyes. Most owls are nocturnal , actively hunting their prey in darkness.

Several types of owls, however, are crepuscular —active during the twilight hours of dawn and dusk; one example is the pygmy owl Glaucidium.

A few owls are active during the day, also; examples are the burrowing owl Speotyto cunicularia and the short-eared owl Asio flammeus. Much of the owls' hunting strategy depends on stealth and surprise.

Owls have at least two adaptations that aid them in achieving stealth. First, the dull coloration of their feathers can render them almost invisible under certain conditions.

Secondly, serrated edges on the leading edge of owls' remiges muffle an owl's wing beats, allowing an owl's flight to be practically silent. Some fish-eating owls, for which silence has no evolutionary advantage, lack this adaptation.

An owl's sharp beak and powerful talons allow it to kill its prey before swallowing it whole if it is not too big. Scientists studying the diets of owls are helped by their habit of regurgitating the indigestible parts of their prey such as bones, scales, and fur in the form of pellets.

These "owl pellets" are plentiful and easy to interpret, and are often sold by companies to schools for dissection by students as a lesson in biology and ecology.

Owl eggs typically have a white colour and an almost spherical shape, and range in number from a few to a dozen, depending on species and the particular season; for most, three or four is the more common number.

In at least one species, female owls do not mate with the same male for a lifetime. Female burrowing owls commonly travel and find other mates, while the male stays in his territory and mates with other females.

The systematic placement of owls is disputed. For example, the Sibley—Ahlquist taxonomy of birds finds that, based on DNA-DNA hybridization , owls are more closely related to the nightjars and their allies Caprimulgiformes than to the diurnal predators in the order Falconiformes ; consequently, the Caprimulgiformes are placed in the Strigiformes, and the owls in general become a family, the Strigidae.

A recent study indicates that the drastic rearrangement of the genome of the accipitrids may have obscured any close relationship of theirs with groups such as the owls.

Some to extant species of owls are known, subdivided into two families: 1. Some entirely extinct families have also been erected based on fossil remains; these differ much from modern owls in being less specialized or specialized in a very different way such as the terrestrial Sophiornithidae.

The Paleocene genera Berruornis and Ogygoptynx show that owls were already present as a distinct lineage some 60—57 million years ago Mya , hence, possibly also some 5 million years earlier, at the extinction of the non-avian dinosaurs.

This makes them one of the oldest known groups of non- Galloanserae landbirds. The supposed " Cretaceous owls" Bradycneme and Heptasteornis are apparently non- avialan maniraptors.

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